Campbell Biology, Canadian Edition, 4th edition

Published by Pearson Canada (December 8, 2023) © 2025

  • Lisa A. Urry Mills College, Oakland, California
  • Michael L. Cain Bowdoin College, Brunswick, Maine
  • Steven A. Wasserman University of California, San Diego
  • Peter V. Minorsky Mercy College, Dobbs Ferry, New York
  • Rebecca Orr Collin College, McKinney, Texas
  • Kerry L Hull Bishop's University, Quebec, Canada
  • Fiona E. Rawle University of Toronto Mississauga, Mississauga, Ontario
  • Dion G. Durnford University of New Brunswick, Fredericton, New Brunswick
  • Christopher D. Moyes Queens University
  • Kevin GE Scott University of Manitoba

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For courses in general biology

Campbell BIOLOGY sets students on the path to success in biology through its clear and engaging narrative, superior skills instruction, innovative use of art and photos, and fully integrated media resources to enhance teaching and learning.

Hallmark features of this title

  • Primary among the themes of Campbell BIOLOGY is evolution. Chapters throughout the text include at least one Evolution section that explicitly focuses on evolutionary aspects of the chapter material, and chapters end with an Evolution Connection Question and a Write About a Theme Question.
  • To help students distinguish the “forest from the trees,” each chapter is organized around a framework of three to seven carefully chosen Key Concepts. The text, Concept Check Questions, Summary of Key Concepts, and MasteringBiology all reinforce these main ideas and essential facts.
  • Campbell BIOLOGY has always featured scientific inquiry, an essential component of any biology course. Complementing stories of scientific discovery in the text narrative, the unit-opening interviews, and our standard-setting Inquiry Figures all deepen the ability of students to understand how we know what we know. Scientific Inquiry Questions give students opportunities to practice scientific thinking, along with the new Scientific Skills Exercises and Interpret the Data Questions. Together, these activities provide students practice both in applying the process of science and in using quantitative reasoning.

New GenAI Study Tool

We’ve heard how important it is for students to use reliable AI tools in responsible and productive ways. To that end, Pearson is focused on creating tools that combine the power of generative AI with trusted Pearson content to provide students with a simplified study experience, delivering on-demand and personalised support that compliments your teaching and aligns with the text you’ve chosen. The Study Tool is available to students who access the Pearson eText on its own or through MyLab.

What can the AI Study Tool do?

  • Generate simplified explanations of challenging sections
  • Summarize material to help learners focus on key topics and ideas
  • Students can ask for multiple choice or short answer questions related to a specific chapter or section to help fill knowledge gaps
  • For extra practice, students can also generate flashcards and notes based on their chat interaction with the tool

New to this Edition

  • As the world shifts to a greater reliance on digital media, it is appropriate that this resource evolves as well. This fourth Canadian edition is the first fully digital version of Campbell Biology. Instructors and students will find that, although the medium has changed, the content is fully consistent with prior editions.
  • In an effort to act on the Calls to Action from the Truth and Reconciliation Commissioners' Report (2012), the Canadian authors were committed to including more Indigenous content in this text, under the guidance and advice of Indigenous students and an Indigenous instructor. Many of the words we use to refer to flora and fauna, as well as locations, (such as caribou, moose, Toronto, Ontario, Saskatchewan, and Canada) all come from Indigenous origins, and this edition acknowledges and highlights these Indigenous word roots. This edition also includes references to how Traditional Ecological Knowledge (TEK) is being used in various contexts throughout the text, especially in Unit 8.
  • Because text and illustrations are equally important for learning biology, integration of text and figures has been a hallmark of this text since the first edition. In addition to the new Visualizing Figures, our popular Exploring Figures and Make Connections Figures epitomize this approach. Each Exploring Figure is a learning unit of core content that brings together related illustrations and text. Make Connections Figures reinforce fundamental conceptual connections throughout biology, helping students overcome tendencies to compartmentalize information. The Fourth Canadian Edition features Guided Tour Figures that walk students through complex figures as an instructor would.

Important Digital Assets in Mastering

  • New GenAI Study Tool. We’ve heard how important it is for students to use reliable AI tools in responsible and productive ways. To that end, Pearson is focused on creating tools that combine the power of generative AI with trusted Pearson content to provide students with a simplified study experience, delivering on-demand and personalised support that compliments your teaching and aligns with the text you’ve chosen.
  • Interactive Figure Walk-through Videos. Figure Walkthroughs guide students through key figures with narrated explanations, figure markups, and questions that reinforce important points. Questions embedded in each Figure Walkthrough encourage students to be active participants in their learning.The videos are in the eText and included in assignable activities in Mastering Biology that allow students to apply what they've learned.
  • Making Models Activities. Whiteboard videos - accessible online via the Study Area in Mastering Biology help students develop their visual modelling skills. The videos are also included in assignable activities that allow students to practise modelling and to apply their understanding to new situations.
  • New! CheckPoint Interactive Questions Embedded at point-of-use in the eText, these short sequences of interactive questions diagnose and remediate misconceptions around topics that are common “stumbling blocks” to understanding. Customized feedback guides students through an evidence-based approach to reason their way to learning and retaining the content. The CheckPoints are also assignable in Mastering Biology.
  • New! Data Modelling Activities. These are simulations that allow students to make predictions, manipulate variables, and examine data.
  • Case Study Questions. Put It All Together Case Study questions are assignable in Mastering Biology. Additional clicker questions are also provided in instructor resources to facilitate classroom activities.
  • Solve It Tutorials. These activities allow students to act like scientists in simulated investigations. Each tutorial presents an interesting, real-world question that students will answer by analyzing and interpreting data.
  • Experimental Inquiry Tutorials. The call to teach students about the process of science has never been louder. To support such teaching, there are 10 interactive tutorials on classic scientific experiments—ranging from Meselson–Stahl on DNA replication to the Grants' work on Galápagos finches and Connell's work on competition. Students who use these tutorials should be better prepared to think critically about experimental design and evaluate the wider implications of the data—preparing them to do the work of real scientists in the future.
  • BioFlix® 3-D Animations. BioFlix® 3-D Animations help students visualize biology with movie-quality animations that can be presented in class, reviewed by students on their own in the Study Area or eText, and assigned in Mastering Biology. BioFlix Tutorials use the animations as a jumping-off point for Mastering Biology coaching exercises on tough topics.
  • HHMI Short Films. Documentary-quality movies from the Howard Hughes Medical Institute that explore topics from the discovery of the double helix to evolution with assignable questions are available in Mastering Biology with assignable questions to make sure students understand key ideas.
  • Galápagos Evolution Video Activities. These video activities, filmed on the Galapagos Islands by Peter and Rosemary Grant, bring to life the dynamic evolutionary processes that impact Darwin's finches on Daphne Major Island. These explore important concepts and data from the Grants' field research and include assignable activities.
  • GraphIt! Coaching Activities. Interpreting Graphs and Data Activities help students develop basic data analysis skills and practise applying these skills by interpreting data about environmental issues.
  • Big Picture Concept Map Tutorials. A new, more engaging concept mapping tool is the basis for highly interactive, challenging concept map activities based on the Big Picture figures in the textbook. Students build their own concept maps, which are auto-graded, and then answer questions to make sure they understand key ideas and make important connections.
  • BioSkills Activities. Activities based on the BioSkills content in the textbook are assignable in Mastering Biology, including activities to support the new BioSkills.
  1. Evolution, the Themes of Biology, and Scientific Inquiry
  2. The Chemical Context of Life
  3. Water and Life
  4. Carbon and the Molecular Diversity of Life
  5. The Structure and Function of Large Biological Molecules
  6. A Tour of the Cell
  7. Membrane Structure and Function
  8. An Introduction to Metabolism
  9. Cellular Respiration and Fermentation
  10. Photosynthesis
  11. Cell Communication
  12. The Cell Cycle
  13. Meiosis and Sexual Life Cycles
  14. Mendel and the Gene Idea
  15. The Chromosomal Basis of Inheritance
  16. The Molecular Basis of Inheritance
  17. From Gene to Protein
  18. Regulation of Gene Expression
  19. Viruses
  20. DNA Tools and Biotechnology
  21. Genomes and Their Evolution
  22. Descent with Modification: A Darwinian View of Life
  23. The Evolution of Populations
  24. The Origin of Species
  25. The History of Life on Earth
  26. Phylogeny and the Tree of Life
  27. Bacteria and Archaea
  28. Protists
  29. Plant Diversity I: How Plants Colonized Land
  30. Plant Diversity II: The Evolution of Seed Plants
  31. Fungi
  32. An Overview of Animal Diversity
  33. An Introduction to Invertebrates
  34. The Origin and Evolution of Vertebrates
  35. Plant Structure, Growth, and Development
  36. Resource Acquisition and Transport in Vascular Plants
  37. Soil and Plant Nutrition
  38. Angiosperm Reproduction and Biotechnology
  39. Plant Responses to Internal and External Signals
  40. Basic Principles of Animal Form and Function
  41. Animal Nutrition
  42. Circulation and Gas Exchange
  43. The Immune System
  44. Osmoregulation and Excretion
  45. Hormones and the Endocrine System
  46. Animal Reproduction
  47. Animal Development
  48. Neurons, Synapses, and Signalling
  49. Nervous Systems
  50. Sensory and Motor Mechanisms
  51. Animal Behaviour
  52. An Introduction to Ecology and the Biosphere
  53. Population Ecology
  54. Community Ecology
  55. Ecosystems and Restoration Ecology
  56. Conservation Biology and Global Change

Lisa A. Urry is Professor of Biology and Chair of the Biology Department at Mills College in Oakland, California, and a Visiting Scholar at the University of California, Berkeley. After graduating from Tufts University with a double major in biology and French, Lisa completed her Ph.D. in molecular and developmental biology at the Massachusetts Institute of Technology (MIT) in the MIT/Woods Hole Oceanographic Institution Joint Program. She has published a number of research papers, most of them focused on gene expression during embryonic and larval development in sea urchins. Lisa has taught a variety of courses, from introductory biology to developmental biology and senior seminar. As a part of her mission to increase understanding of evolution, Lisa also teaches a nonmajors course called Evolution for Future Presidents and is on the Teacher Advisory Board for the Understanding Evolution website developed by the University of California Museum of Paleontology. Lisa is also deeply committed to promoting opportunities in science for women and underrepresented minorities. Lisa is also a co-author of Campbell Biology in Focus.

Michael L. Cain is an ecologist and evolutionary biologist who is now writing full-time. Michael earned a joint degree in biology and math at Bowdoin College, an M.Sc. from Brown University, and a Ph.D. in ecology and evolutionary biology from Cornell University. As a faculty member at New Mexico State University and Rose-Hulman Institute of Technology, he taught a wide range of courses, including introductory biology, ecology, evolution, botany, and conservation biology. Michael is the author of dozens of scientific papers on topics that include foraging behaviour in insects and plants, long-distance seed dispersal, and speciation in crickets. In addition to his work on Campbell BIOLOGY and Campbell Biology in Focus, Michael is the lead author of an ecology textbook.

Steven A. Wasserman is Professor of Biology at the University of California, San Diego (UCSD). He earned his A.B. in biology from Harvard University and his Ph.D. in biological sciences from MIT. through his research on regulatory pathway mechanisms in the fruit fly Drosophila, Steve has contributed to the fields of developmental biology, reproduction, and immunity. As a faculty member at the University of Texas Southwestern Medical Center and UCSD, he has taught genetics, development, and physiology to undergraduate, graduate, and medical students. He currently focuses on teaching introductory biology. He has also served as the research mentor for more than a dozen doctoral students and more than 50 aspiring scientists at the undergraduate and high school levels. Steve has been the recipient of distinguished scholar awards from both the Markey Charitable Trust and the David and Lucile Packard Foundation. In 2007, he received UCSD's Distinguished Teaching Award for undergraduate teaching. Steve is also a co-author of Campbell Biology in Focus. 

Peter V. Minorsky is Professor of Biology at Mercy College in New York, where he teaches introductory biology, evolution, ecology, and botany. He received his A.B. in biology from Vassar College and his Ph.D. in plant physiology from Cornell University. He is also the science writer for the journal Plant Physiology. After a postdoctoral fellowship at the University of Wisconsin at Madison, Peter taught at Kenyon College, Union College, Western Connecticut State University, and Vassar College. His research interests concern how plants sense environmental change. Peter received the 2008 Award for Teaching Excellence at Mercy College. Peter is also a co-author of Campbell Biology in Focus.

Rebecca B. Orr (Ready-to-Go Teaching Modules, Interactive Visual Activities, eText Media Integration) is Professor of Biology at Collin College in Plano, Texas, where she teaches introductory biology. She earned her B.S. from Texas A&M University and her Ph.D. from University of Texas Southwestern Medical Center at Dallas. Rebecca has a passion for investigating strategies that result in more effective learning and retention, and she is a certified Team-Based Learning Collaborative Trainer Consultant. She enjoys focusing on the creation of learning opportunities that both engage and challenge students.

Fiona Rawle received her Ph.D. from Queen's University in Kingston, Ontario. She is a Professor, Teaching Stream, at the University of Toronto Mississauga, where she teaches Introduction to Evolution and Evolutionary Genetics, Introductory Genetics, and Molecular Basis of Disease. Fiona's teaching and pedagogical research interests focus on several areas: (1) learning from failure in science; (2) science misinformation and scientific literacy; (3) bias in STEAM; and (4) the development of active learning techniques that can be used in large class settings. Fiona was the recipient of a 3M National Teaching Fellowship in 2022, Canada's highest honour for teaching in higher education.

Dion Durnford is a professor at the University of New Brunswick, in Fredericton. He earned a B.Sc. in Biology from Dalhousie University and a Ph.D. in Botany from the University of British Columbia. His research has focused on the evolution of light-harvesting antenna systems and the role of these proteins in light harvesting and photo-protection in microalgae. His recent work is examining how microalgae age and their strategies for increasing longevity. Dion was the recipient of the 2002 Faculty of Science Excellence in Teaching award and the 2010 Allan P. Stewart Award for Excellence in Teaching.

Chris Moyes is a comparative physiologist, focusing on the muscle biochemistry and energetics. He received his Ph.D. in Zoology from the University of British Columbia (1991) and is currently a Professor in the Department of Biology, Queen's University. He has published more than 100 research papers and contributed to four books. He is co-author of Principles of Animal Physiology, first published in 2006.

Kevin Scott is a senior instructor at the University of Manitoba where he teaches introductory biology for both biology majors and nonbiology majors; human physiology; and environmental physiology of animal laboratories. In the past, he has also taught courses in ecology for nonbiology majors, immunology, parasitology, and microbiology. He received a B.Sc. in Zoology and a Ph.D. joint between Zoology and Cellular, Molecular, and Microbial Biology at the University of Calgary. As an educator, Dr. Scott's career is centred on teaching and the classroom, where he shares his excitement for biology. His interest in plant biology has grown during his professional career and is a favourite topic in his classroom. Kevin was a co-author of Campbell Biology: Concepts and Connections, Canadian Edition.

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