Table of contents
- 1. Introduction to Biology2h 42m
- 2. Chemistry3h 40m
- 3. Water1h 26m
- 4. Biomolecules2h 23m
- 5. Cell Components2h 26m
- 6. The Membrane2h 31m
- 7. Energy and Metabolism2h 0m
- 8. Respiration2h 40m
- 9. Photosynthesis2h 49m
- 10. Cell Signaling59m
- 11. Cell Division2h 47m
- 12. Meiosis2h 0m
- 13. Mendelian Genetics4h 44m
- Introduction to Mendel's Experiments7m
- Genotype vs. Phenotype17m
- Punnett Squares13m
- Mendel's Experiments26m
- Mendel's Laws18m
- Monohybrid Crosses19m
- Test Crosses14m
- Dihybrid Crosses20m
- Punnett Square Probability26m
- Incomplete Dominance vs. Codominance20m
- Epistasis7m
- Non-Mendelian Genetics12m
- Pedigrees6m
- Autosomal Inheritance21m
- Sex-Linked Inheritance43m
- X-Inactivation9m
- 14. DNA Synthesis2h 27m
- 15. Gene Expression3h 20m
- 16. Regulation of Expression3h 31m
- Introduction to Regulation of Gene Expression13m
- Prokaryotic Gene Regulation via Operons27m
- The Lac Operon21m
- Glucose's Impact on Lac Operon25m
- The Trp Operon20m
- Review of the Lac Operon & Trp Operon11m
- Introduction to Eukaryotic Gene Regulation9m
- Eukaryotic Chromatin Modifications16m
- Eukaryotic Transcriptional Control22m
- Eukaryotic Post-Transcriptional Regulation28m
- Eukaryotic Post-Translational Regulation13m
- 17. Viruses37m
- 18. Biotechnology2h 58m
- 19. Genomics17m
- 20. Development1h 5m
- 21. Evolution3h 1m
- 22. Evolution of Populations3h 52m
- 23. Speciation1h 37m
- 24. History of Life on Earth2h 6m
- 25. Phylogeny2h 31m
- 26. Prokaryotes4h 59m
- 27. Protists1h 12m
- 28. Plants1h 22m
- 29. Fungi36m
- 30. Overview of Animals34m
- 31. Invertebrates1h 2m
- 32. Vertebrates50m
- 33. Plant Anatomy1h 3m
- 34. Vascular Plant Transport1h 2m
- 35. Soil37m
- 36. Plant Reproduction47m
- 37. Plant Sensation and Response1h 9m
- 38. Animal Form and Function1h 19m
- 39. Digestive System1h 10m
- 40. Circulatory System1h 57m
- 41. Immune System1h 12m
- 42. Osmoregulation and Excretion50m
- 43. Endocrine System1h 4m
- 44. Animal Reproduction1h 2m
- 45. Nervous System1h 55m
- 46. Sensory Systems46m
- 47. Muscle Systems23m
- 48. Ecology3h 11m
- Introduction to Ecology20m
- Biogeography14m
- Earth's Climate Patterns50m
- Introduction to Terrestrial Biomes10m
- Terrestrial Biomes: Near Equator13m
- Terrestrial Biomes: Temperate Regions10m
- Terrestrial Biomes: Northern Regions15m
- Introduction to Aquatic Biomes27m
- Freshwater Aquatic Biomes14m
- Marine Aquatic Biomes13m
- 49. Animal Behavior28m
- 50. Population Ecology3h 41m
- Introduction to Population Ecology28m
- Population Sampling Methods23m
- Life History12m
- Population Demography17m
- Factors Limiting Population Growth14m
- Introduction to Population Growth Models22m
- Linear Population Growth6m
- Exponential Population Growth29m
- Logistic Population Growth32m
- r/K Selection10m
- The Human Population22m
- 51. Community Ecology2h 46m
- Introduction to Community Ecology2m
- Introduction to Community Interactions9m
- Community Interactions: Competition (-/-)38m
- Community Interactions: Exploitation (+/-)23m
- Community Interactions: Mutualism (+/+) & Commensalism (+/0)9m
- Community Structure35m
- Community Dynamics26m
- Geographic Impact on Communities21m
- 52. Ecosystems2h 36m
- 53. Conservation Biology24m
49. Animal Behavior
Animal Behavior
Problem 2`
Textbook Question
According to Hamilton's rule,
a. Natural selection does not favor altruistic behavior that causes the death of the altruist.
b. Natural selection favors altruistic acts when the resulting benefit to the recipient, corrected for relatedness, exceeds the cost to the altruist.
c. Natural selection is more likely to favor altruistic behavior that benefits an offspring than altruistic behavior that benefits a sibling.
d. The effects of kin selection are larger than the effects of direct natural selection on individuals.

1
Understand Hamilton's rule, which is a principle in evolutionary biology that explains when altruistic behavior can evolve through natural selection.
Hamilton's rule is expressed mathematically as \( rB > C \), where \( r \) is the genetic relatedness between the altruist and the recipient, \( B \) is the benefit to the recipient, and \( C \) is the cost to the altruist.
Analyze each option in the problem statement to determine which one aligns with Hamilton's rule. Focus on the relationship between the benefit to the recipient, the cost to the altruist, and the genetic relatedness.
Option b states that natural selection favors altruistic acts when the resulting benefit to the recipient, corrected for relatedness, exceeds the cost to the altruist. This directly corresponds to Hamilton's rule \( rB > C \).
Review the other options to ensure they do not align with Hamilton's rule. Option a, c, and d do not accurately reflect the mathematical relationship described by Hamilton's rule.

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Key Concepts
Here are the essential concepts you must grasp in order to answer the question correctly.
Hamilton's Rule
Hamilton's Rule is a principle in evolutionary biology that predicts when altruistic behavior will evolve. It states that natural selection favors altruistic acts if the benefit to the recipient, multiplied by the coefficient of relatedness, exceeds the cost to the altruist. This rule is often summarized by the inequality: rB > C, where r is relatedness, B is the benefit to the recipient, and C is the cost to the altruist.
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Chargaff's Rules
Altruistic Behavior
Altruistic behavior in biology refers to actions by an individual that benefit another individual at a cost to itself. This behavior is puzzling from an evolutionary perspective because it seems to reduce the altruist's fitness. However, through mechanisms like kin selection, altruism can be advantageous if it increases the survival and reproductive success of related individuals who share genes with the altruist.
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Migration and Altruism
Kin Selection
Kin selection is a form of natural selection that considers the role of relatives in the evolution of social behaviors. It suggests that individuals can increase their genetic success through behaviors that help relatives, even at a personal cost. This concept helps explain altruistic behaviors, as aiding relatives can ensure the propagation of shared genes, thus indirectly benefiting the altruist's genetic lineage.
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